Many new species of Okenia

December 21, 2004
From: Bill Rudman

In the last 3 months four papers have been published describing many new species of the goniodorid genus Okenia from the western and eastern Pacific. Also included is further information on the anatomy and biology on many already named species. One paper (Gosliner, 2004) is causing particular angst because it proposes discarding the genus Hopkinsia, a name dear to the hearts of the 'branchers' of California. In the same review other genera such as Hopkinsiella and Sakishimaia are also shown to be unnecessary. I have followed Gosliner's proposal on the Forum. New Fact Sheets have been prepared for all the new species, and many existing Fact Sheets have been revised to incorporate the new information now available to us. As Fact Sheets are often cited in scientific publications, I am keeping copies of all replaced Fact Sheets as pdf files, which will be available on request if anyone has a need to refer to them.

Listed below are the new Fact Sheets and messages, and the relevant publications.

Okenia brunneomaculata Gosliner, 2004
Okenia purpureolineata Gosliner, 2004
Okenia kendi Gosliner, 2004
Okenia virginiae Gosliner, 2004
Okenia lambat Gosliner, 2004
Okenia liklik Gosliner, 2004
More information is provided on: 
Okenia pellucida Burn, 1967
Okenia japonica   Baba, 1949
Okenia rosacea  (MacFarland, 1905) [formerly Hopkinsia]
Okenia plana Baba, 1960 [formerly Hopkinsia]
Okenia pilosa (Bouchet and Ortea, 1983) [formerly Hopkinsia]
Okenia nakamotoensis (Hamatani, 2001) [formerly Hopkinsia]
Okenia kondoi (Hamatani, 2001) [formerly Sakishimaia]
[In:] Gosliner, T. M. (2004)
Phylogenetic Systematics of Okenia, Sakishimaia, Hopkinsiella and Hopkinsia (Nudibranchia: Goniodorididae) with descriptions of new species from the tropical Indo-Pacific. Proceedings of the California Academy of Sciences, 55: 125-161.

Okenia purpurata Rudman, 2004
Okenia vena Rudman, 2004
Okenia mellita Rudman, 2004
Okenia stellata Rudman, 2004
Okenia hallucigenia Rudman, 2004
More information is provided on:
Okenia barnardi Baba, 1937.
Okenia pellucida Burn, 1967
Okenia japonica Baba, 1949
Okenia mija Burn, 1967
Okenia zoobotryon (Smallwood, 1910)
Okenia hiroi (Baba, 1938) [formerly Hopkinsia]
Okenia pilosa (Bouchet and Ortea, 1983) [formerly Hopkinsia]
Okenia plana Baba, 1960 [formerly Hopkinsia]
Okenia echinata Baba, 1949
Okenia virginiae Gosliner, 2004
[In:] Rudman, W.B. (2004) Further species of the opisthobranch genus Okenia (Nudibranchia: Goniodorididae) from the Indo-West Pacific. Zootaxa, 695: 1-70.

Okenia angelica Gosliner & Bertsch,  2004 
Okenia cochimi Gosliner & Bertsch, 2004
Okenia mexicorum Gosliner & Bertsch, 2004
More information is provided on:
Okenia vancouverensis (O'Donoghue, 1921)
Okenia angelensis Lance, 1966
[In:] Gosliner, T. M. & Bertsch, H. (2004) Systematics of Okenia from the Pacific Coast of North America (Nudibranchia: Goniodorididae) with descriptions of three new species. Proceedings of the California Academy of Sciences, 55: 412-428, figs. 1-11.

Okenia academica Camacho-García & Gosliner, 2004
[In:] Camacho-García, Y.E. & Gosliner, T.M. (2004) A New Species of Okenia (Gastropoda: Nudibranchia: Goniodorididae) from the Pacific Coast of Costa Rica. Proceedings of the California Academy of Sciences, 55: 431-438.

Bill Rudman

Rudman, W.B., 2004 (Dec 21) Many new species of Okenia. [Message in] Sea Slug Forum. Australian Museum, Sydney. Available from


Okenia rosacea

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